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Beetles with somewhat flattened, highly mobile larvae include the ground beetles and rove beetles; their larvae are described as campodeiform.

Some beetle larvae resemble hardened worms with dark head capsules and minute legs. These are elateriform larvae, and are found in the click beetle Elateridae and darkling beetle Tenebrionidae families.

Some elateriform larvae of click beetles are known as wireworms. Beetles in the Scarabaeoidea have short, thick larvae described as scarabaeiform, more commonly known as grubs.

All beetle larvae go through several instars , which are the developmental stages between each moult. In many species, the larvae simply increase in size with each successive instar as more food is consumed.

In some cases, however, more dramatic changes occur. Among certain beetle families or genera, particularly those that exhibit parasitic lifestyles, the first instar the planidium is highly mobile to search out a host, while the following instars are more sedentary and remain on or within their host.

This is known as hypermetamorphosis ; it occurs in the Meloidae , Micromalthidae , and Ripiphoridae. Its first stage, the triungulin , has longer legs to go in search of the eggs of grasshoppers.

After feeding for a week it moults to the second stage, called the caraboid stage, which resembles the larva of a carabid beetle.

In another week it moults and assumes the appearance of a scarabaeid larva — the scarabaeidoid stage. Its penultimate larval stage is the pseudo-pupa or the coarcate larva, which will overwinter and pupate until the next spring.

The larval period can vary widely. A fungus feeding staphylinid Phanerota fasciata undergoes three moults in 3. Leiodidae completes its larval stage in the fruiting body of slime mold in 2 days and possibly represents the fastest growing beetles.

Dermestid beetles, Trogoderma inclusum can remain in an extended larval state under unfavourable conditions, even reducing their size between moults.

A larva is reported to have survived for 3. As with all endopterygotes, beetle larvae pupate, and from these pupae emerge fully formed, sexually mature adult beetles, or imagos.

Pupae never have mandibles they are adecticous. In most pupae, the appendages are not attached to the body and are said to be exarate ; in a few beetles Staphylinidae, Ptiliidae etc.

Adults have extremely variable lifespans, from weeks to years, depending on the species. It is believed that when furniture or house timbers are infested by beetle larvae, the timber already contained the larvae when it was first sawn up.

A birch bookcase 40 years old released adult Eburia quadrigeminata Cerambycidae , while Buprestis aurulenta and other Buprestidae have been documented as emerging as much as 51 years after manufacture of wooden items.

The elytra allow beetles to both fly and move through confined spaces, doing so by folding the delicate wings under the elytra while not flying, and folding their wings out just before take off.

The unfolding and folding of the wings is operated by muscles attached to the wing base; as long as the tension on the radial and cubital veins remains, the wings remain straight.

In some day-flying species for example, Buprestidae , Scarabaeidae , flight does not include large amounts of lifting of the elytra, having the metathorac wings extended under the lateral elytra margins.

Many rove beetles have greatly reduced elytra, and while they are capable of flight, they most often move on the ground: Diving beetles Dytiscidae hold air between the abdomen and the elytra when diving.

Hydrophilidae have hairs on their under surface that retain a layer of air against their bodies. Adult crawling water beetles use both their elytra and their hind coxae the basal segment of the back legs in air retention, while whirligig beetles simply carry an air bubble down with them whenever they dive.

Beetles have a variety of ways to communicate, including the use of pheromones. The mountain pine beetle emits a pheromone to attract other beetles to a tree.

The mass of beetles are able to overcome the chemical defenses of the tree. This species can stridulate to communicate [88] , but others may use sound to defend themselves when attacked [89].

Parental care is found in a few species of beetle, perhaps for protection against adverse conditions and predators.

Burying beetles are attentive parents, and participate in cooperative care and feeding of their offspring. Both parents work to bury small animal carcass to serve as a food resource for their young and build a brood chamber around it.

The parents prepare the carcass and protect it from competitors and from early decomposition. After their eggs hatch, the parents keep the larvae clean of fungus and bacteria and help the larvae feed by regurgitating food for them.

Some dung beetles provide parental care, collecting herbivore dung and laying eggs within that food supply, an instance of mass provisioning.

Some species do not leave after this stage, but remain to safeguard their offspring. Most species of beetles do not display parental care behaviors after the eggs have been laid.

Eusociality involves cooperative brood care including brood care of offspring from other individuals , overlapping generations within a colony of adults, and a division of labour into reproductive and non-reproductive groups.

It is one of more than species of wood-boring Ambrosia beetles which distribute the spores of ambrosia fungi. Beetles are able to exploit a wide diversity of food sources available in their many habitats.

Some are omnivores , eating both plants and animals. Other beetles are highly specialized in their diet. Many species of leaf beetles, longhorn beetles, and weevils are very host-specific, feeding on only a single species of plant.

Ground beetles and rove beetles Staphylinidae , among others, are primarily carnivorous and catch and consume many other arthropods and small prey, such as earthworms and snails.

While most predatory beetles are generalists, a few species have more specific prey requirements or preferences.

Decaying organic matter is a primary diet for many species. This can range from dung , which is consumed by coprophagous species such as certain scarab beetles in the Scarabaeidae , to dead animals, which are eaten by necrophagous species such as the carrion beetles , Silphidae.

Some beetles found in dung and carrion are in fact predatory. These include members of the Histeridae and Silphidae , preying on the larvae of coprophagous and necrophagous insects.

Some beetles have special mycangia , structures for the transport of fungal spores. Beetles, both adults and larvae, are the prey of many animal predators including mammals from bats to rodents , birds , lizards , amphibians , fishes , dragonflies , robberflies , reduviid bugs , ants , other beetles, and spiders.

These include camouflage and mimicry against predators that hunt by sight, toxicity, and defensive behaviour.

Camouflage is common and widespread among beetle families, especially those that feed on wood or vegetation, such as leaf beetles Chrysomelidae, which are often green and weevils.

In some species, sculpturing or various coloured scales or hairs cause beetles such as the avocado weevil Heilipus apiatus to resemble bird dung or other inedible objects.

Some longhorn beetles Cerambycidae are effective Batesian mimics of wasps. Beetles may combine coloration with behavioural mimicry, acting like the wasps they already closely resemble.

Many other beetles, including ladybirds , blister beetles , and lycid beetles secrete distasteful or toxic substances to make them unpalatable or poisonous, and are often aposematic , where bright or contrasting coloration warn off predators; many beetles and other insects mimic these chemically protected species.

Chemical defense is important in some species, usually being advertised by bright aposematic colours. Some Tenebrionidae use their posture for releasing noxious chemicals to warn off predators.

Chemical defences may serve purposes other than just protection from vertebrates, such as protection from a wide range of microbes. Some species sequester chemicals from the plants they feed on, incorporating them into their own defenses.

Other species have special glands to produce deterrent chemicals. The defensive glands of carabid ground beetles produce a variety of hydrocarbons , aldehydes , phenols , quinones , esters , and acids released from an opening at the end of the abdomen.

African carabid beetles for example, Anthia and Thermophilum — Thermophilum is sometimes included within Anthia employ the same chemicals as ants: The gland is made of two containing chambers, one for hydroquinones and hydrogen peroxide , the other holding hydrogen peroxide and catalase enzymes.

The oxygen propels the noxious chemical spray as a jet that can be aimed accurately at predators. Large ground-dwelling beetles such as Carabidae , the rhinoceros beetle and the longhorn beetles defend themselves using strong mandibles , or heavily sclerotised armored spines or horns to deter or fight off predators.

Some combine it with thanatosis , in which they close up their appendages and "play dead". A few species of beetles are ectoparasitic on mammals.

One such species, Platypsyllus castoris , parasitises beavers Castor spp. This beetle lives as a parasite both as a larva and as an adult, feeding on epidermal tissue and possibly on skin secretions and wound exudates.

They are wingless and eyeless, as are many other ectoparasites. Beetle-pollinated flowers are usually large, greenish or off-white in color, and heavily scented.

Scents may be spicy, fruity, or similar to decaying organic material. Beetles were most likely the first insects to pollinate flowers. Most beetle-pollinated flowers are flattened or dish-shaped, with pollen easily accessible, although they may include traps to keep the beetle longer.

The beetle families that habitually pollinate flowers are the Buprestidae, Cantharidae, Carambycidae, Cleridae, Dermestidae, Lycidae, Melyridae, Mordellidae, Nitidulidae and Scarabeidae.

Mutualism is well known in a few beetles, such as the ambrosia beetle , which partners with fungi to digest the wood of dead trees.

The beetles excavate tunnels in dead trees in which they cultivate fungal gardens, their sole source of nutrition.

After landing on a suitable tree, an ambrosia beetle excavates a tunnel in which it releases spores of its fungal symbiont. The beetles cannot eat the wood due to toxins, and uses its relationship with fungi to help overcome the defenses of its host tree in order to provide nutrition for their larvae.

Adult diapause is the most common form of diapause in Coleoptera. To endure the period without food often lasting many months adults prepare by accumulating reserves of lipids, glycogen, proteins and other substances needed for resistance to future hazardous changes of environmental conditions.

This diapause is induced by signals heralding the arrival of the unfavourable season; usually the cue is photoperiodic.

Short decreasing day length serves as a signal of approaching winter and induces winter diapause hibernation. This loss of body fat was a gradual process, occurring in combination with dehydration.

All insects are poikilothermic , [] so the ability of a few beetles to live in extreme environments depends on their resilience to unusually high or low temperatures.

The low temperatures experienced by Cucujus clavipes can be survived through their deliberate dehydration in conjunction with the antifreeze proteins.

This concentrates the antifreezes several fold. Conversely, desert dwelling beetles are adapted to tolerate high temperatures.

These beetles also exhibits behavioural adaptions to tolerate the heat: The fogstand beetle of the Namib Desert , Stenocara gracilipes , is able to collect water from fog , as its elytra have a textured surface combining hydrophilic water-loving bumps and waxy, hydrophobic troughs.

The beetle faces the early morning breeze, holding up its abdomen; droplets condense on the elytra and run along ridges towards their mouthparts.

Similar adaptations are found in several other Namib desert beetles such as Onymacris unguicularis. Some terrestrial beetles that exploit shoreline and floodplain habitats have physiological adaptations for surviving floods.

In the event of flooding, adult beetles may be mobile enough to move away from flooding, but larvae and pupa often cannot. Adults of Cicindela togata are unable to survive immersion in water, but larvae are able to survive a prolonged period, up to 6 days, of anoxia during floods.

Anoxia tolerance in the larvae may have been sustained by switching to anaerobic metabolic pathways or by reducing metabolic rate. Many beetle species undertake annual mass movements which are termed as migrations.

These include the pollen beetle Meligethes aeneus [] and many species of coccinellids. Several species of dung beetle, especially the sacred scarab, Scarabaeus sacer , were revered in Ancient Egypt.

The scarab was of prime significance in the funerary cult of ancient Egypt. The best-known of these are the Judean LMLK seals , where eight of 21 designs contained scarab beetles, which were used exclusively to stamp impressions on storage jars during the reign of Hezekiah.

Pliny the Elder discusses beetles in his Natural History , [] describing the stag beetle: To these insects a sting has been denied by Nature; but in one large kind we find horns of a remarkable length, two-pronged at the extremities, and forming pincers, which the animal closes when it is its intention to bite.

He is black, long and has hard wings like a great dung beetle". Many feed on economically important plants and stored plant products, including trees, cereals, tobacco, and dried fruits.

The boll weevil crossed the Rio Grande near Brownsville , Texas, to enter the United States from Mexico around , [] and had reached southeastern Alabama by It remains the most destructive cotton pest in North America.

The bark beetle, elm leaf beetle and the Asian longhorned beetle Anoplophora glabripennis [] are among the species that attack elm trees.

Bark beetles Scolytidae carry Dutch elm disease as they move from infected breeding sites to healthy trees.

The disease has devastated elm trees across Europe and North America. Some species of beetle have evolved immunity to insecticides. For example, the Colorado potato beetle , Leptinotarsa decemlineata , is a destructive pest of potato plants.

Its hosts include other members of the Solanaceae , such as nightshade , tomato , eggplant and capsicum , as well as the potato.

Different populations have between them developed resistance to all major classes of insecticide. The death watch beetle , Xestobium rufovillosum Ptinidae , is a serious pest of older wooden buildings in Europe.

It attacks hardwoods such as oak and chestnut , always where some fungal decay has taken or is taking place. The actual introduction of the pest into buildings is thought to take place at the time of construction.

Other pests include the coconut hispine beetle, Brontispa longissima , which feeds on young leaves , seedlings and mature coconut trees, causing serious economic damage in the Philippines.

Beetles can be beneficial to human economics by controlling the populations of pests. The larvae and adults of some species of lady beetles Coccinellidae feed on aphids that are pests.

Other lady beetles feed on scale insects , whitefly and mealybugs. For example, the genus Zygogramma is native to North America but has been used to control Parthenium hysterophorus in India and Ambrosia artemisiifolia in Russia.

Dung beetles Scarabidae have been successfully used to reduce the populations of pestilent flies, such as Musca vetustissima and Haematobia exigua which are serious pests of cattle in Australia.

The Dermestidae are often used in taxidermy and in the preparation of scientific specimens, to clean soft tissue from bones.

Beetles are the most widely eaten insects, with about species used as food, usually at the larval stage. Due to their habitat specificity, many species of beetles have been suggested as suitable as indicators, their presence, numbers, or absence providing a measure of habitat quality.

Predatory beetles such as the tiger beetles Cicindelidae have found scientific use as an indicator taxon for measuring regional patterns of biodiversity.

They are suitable for this as their taxonomy is stable; their life history is well described; they are large and simple to observe when visiting a site; they occur around the world in many habitats, with species specialised to particular habitats; and their occurrence by species accurately indicates other species, both vertebrate and invertebrate.

Many beetles have beautiful and durable elytra that have been used as material in arts, with beetlewing the best example. Whole beetles, either as-is or encased in clear plastic, are made into objects ranging from cheap souvenirs such as key chains to expensive fine-art jewellery.

In parts of Mexico, beetles of the genus Zopherus are made into living brooches by attaching costume jewelry and golden chains, which is made possible by the incredibly hard elytra and sedentary habits of the genus.

Fighting beetles are used for entertainment and gambling. This sport exploits the territorial behavior and mating competition of certain species of large beetles.

In the Chiang Mai district of northern Thailand, male Xylotrupes rhinoceros beetles are caught in the wild and trained for fighting.

Females are held inside a log to stimulate the fighting males with their pheromones. Beetles are sometimes used as instruments: Some species of beetle are kept as pets , for example diving beetles Dytiscidae may be kept in a domestic fresh water tank.

In Japan the practice of keeping horned rhinoceros beetles Dynastinae and stag beetles Lucanidae is particularly popular amongst young boys.

Beetle collecting became extremely popular in the Victorian era. Several coleopteran adaptations have attracted interest in biomimetics with possible commercial applications.

Living beetles have been used as cyborgs. A Defense Advanced Research Projects Agency funded project implanted electrodes into Mecynorhina torquata beetles, allowing them to be remotely controlled via a radio receiver held on its back, as proof-of-concept for surveillance work.

Many species of beetles have very specific habitats and long life cycles that make them vulnerable. Some species are highly threatened while others are already feared extinct.

In Japan the Genji firefly, Luciola cruciata , is extremely popular, and in South Africa the Addo elephant dung beetle offers promise for broadening ecotourism beyond the big five tourist mammal species.

Popular dislike of pest beetles, too, can be turned into public interest in insects, as can unusual ecological adaptations of species like the fairy shrimp hunting beetle, Cicinis bruchi.

From Wikipedia, the free encyclopedia. This article is about the insect. For the car, see Volkswagen Beetle.

For other uses, see Beetle disambiguation. Insect thermoregulation and Insect winter ecology. Beetlewing and Live insect jewelry.

Retrieved 20 February Species Diversity in Space and Time. An Outline of Entomology 5 ed. Book of Insect Records.

Archived from the original on July 18, New record and remeasuring of Scydosella musawasensis Hall, Coleoptera, Ptiliidae , the smallest known free-living insect".

The beetle fauna of Germany. Retrieved March 16, Evolution of the Insects. Triassic Curculionoidea have the same status as Triassic Chrysoleloidea: Journal of the Kansas Entomological Society.

Archived from the original PDF on July 18, Archived from the original PDF on November 11, Archived from the original PDF on March 31, Archived from the original PDF on July 4, Annalen des Naturhistorischen Museums in Wien.

Archived from the original PDF on The origin of coprophagy ". Journal of Natural History. Proceedings of the Royal Society B: Evolution, Biogeography, and Ecology".

Annual Review of Ecology and Systematics. Tree of Life web project. Retrieved 27 February Handbook of Zoology, Band 4: Morphology, Classification and Keys.

A new beetle family to the fauna of Serbia". Bulletin of the Natural History Museum 6: Carabidae and Trachypachidae ".

Tree of Life Web Project. Retrieved March 18, Retrieved 15 March Journal of Asia-Pacific Entomology. A Guide to the Beetles of Australia.

American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Head, Thorax, Abdomen, and Genitalia Insects ".

Journal of the Royal Society Interface. University of Minnesota Extension. Insects, their structure and life. Entomologische Berichten in Dutch.

The Journal of Experimental Biology. Adaptation and Environment 5th ed. Structure and Physiology of the Circulatory System".

Comprehensive Insect Physiology, Biochemistry and Pharmacology. Integument, Respiration and Circulation. Form, function, and diversity.

Turkish Journal of Zoology. The Biology of the Coleoptera. Medical and Veterinary Entomology 2nd ed. Retrieved 17 March Insights from Vertical-Looking Entomological Radar".

Encyclopedia of Entomology 2 ed. American Beetles, Volume 1. Retrieved March 13, Journal of the New York Entomological Society. The Australian beetle that behaves like a bee".

Beetle Biology And Ecology. Beetles Coleoptera and Coleopterologist. Jahangirnagar University Journal of Biological Science.

Adaptive Coloration in Animals. Gaps in our knowledge of an invasive species". North Carolina State University.

Taxonomy, Ecology and Pathology. Adult Diapause in Coleoptera". A Journal of Entomology. An Outline of Entomology.

Journal of Experimental Biology. Surviving in hostile habitats". The yellow mealworm beetle, Tenebrio molitor, contains a family of small Cys-rich and Thr-rich thermal hysteresis proteins that depress the hemolymph freezing point below the melting point by as much as 5.

Thermal hysteresis protein expression was evaluated throughout development and after exposure to altered environmental conditions.

Journal of Biological Chemistry. Annals of the Entomological Society of America. Wyatt; Stanley, David W. The American Midland Naturalist. The Biology of Life on the Move.

Retrieved 10 February Retrieved July 19, The New Archaeological Excavations at Lachish — Institute of Archaeology of Tel Aviv University.

University of Chicago Press. Insects and other Invertebrates in Classical Antiquity. The Metamorphoses of Antoninus Liberalis: A translation with a commentary.

London and New York, Routledge. Economic impacts of the boll weevil. Archived from the original on May 12, Canadian Journal of Plant Pathology.

American Journal of Potato Research. Why our next terrorist attack could come on six legs". Pest Management and Phytosanitary Trade Barriers. Conservation and collective care.

Archived from the original on July 10, Retrieved July 17, Chrysomelidae in the field". Archived from the original on April 19, Retrieved June 24, Retrieved June 17, Agriculture, Ecosystems and Environment.

Retrieved 14 March Coleoptera, Chrysomelidae in biological control of ragweeds Ambrosia artemisiifolia L. Entomologicheskoe Obozrenije in Russian.

Australian Journal of Entomology. Scarabaeidae can improve soil hydrological properties". In Huffman, Jane E.

Developments in Forensic Science. Preparation protocols for skeletons and fossil vertebrates under the scanning electron microscopy".

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